Christensen, L. B. Marine mammal populations: reconstructing historical abundances at the global scale. Fish. Cent. Res. Rep. 14, 1–161 (2006).
Charpentier, A. et al. What’s in a whale bone? combining new analytical methods, ecology and history to shed light on ancient human-whale interactions. Quat. Sci. Rev. 284, 107470 (2022).
Smith, A. B., Kinahan, J. The invisible whale. World Archaeol. 16, 89–97 (1984).
Gusinde, M. et al. Los Indios de Tierra del Fuego: los Yamana, Buenos Aires, CAEA. (1986).
Turner, L. M. et al. An Aleutian Ethnography, Fairbanks, University of Alaska Press (2008).
Lee, S. M. et al. Chasseurs de baleines: la frise de Bangudae, Corée du Sud, Paris, Errance. (2011).
van den Hurk, Y. et al. The prelude to industrial whaling: identifying the targets of ancient European whaling using zooarchaeology and collagen mass-peptide fingerprinting. R. Soc. Open Sci. 10, 230741 (2023).
Nabais, M., Soares, R. & van den Hurk, Y. The Zooarchaeology of ancient whaling practices in Portugal: A review and a new Roman Republican contribution at Castelo Velho de Safara. PLoS ONE 19, e0310215 (2024).
Erlandson, J. M. The archaeology of aquatic adaptations: paradigms for a new millennium. J. Archaeological Res. 9, 287–350 (2001).
Pétillon, J.-M. et al. Life on the shore of the Bay of Biscay in the Late Upper Paleolithic: towards a new paradigm. In: Dupont, C., Marchand, G. (eds.), Archéologie des chasseurs-cueilleurs maritimes. De la fonction des habitats à l’organisation de l’espace littoral, Paris, Société préhistorique française, 23–36 (2016).
Álvarez-Fernández, E. et al. Mise à jour des recherches sur les chasseurs-cueilleurs côtiers dans le sud-est de la péninsule ibérique durant le Magdalénien: les restes archéofauniques d’origine marine. In: Costamagno, S., Boudadi-Maligne, M., Daujeard, C., Fernandez, P., Stoetzel, E. (eds.), Sociétés humaines et environnements dans la zone circumméditerranéenne du Pléistocène au début de l’Holocène, Les Eyzies-de-Tayac-Sireuil, Musée national de Préhistoire (Paléo hors-série), 50–63 (2022).
Castaños, P. et al. Estudio de los macromamíferos del yacimiento de Santa Catalina. In: Berganza E., Arribas J.L. (eds.), La Cueva de Santa Catalina (Lekeitio): La intervención arqueológica. Restos vegetales, animales y humanos, Bilbao, Kobie serie BAI 4, 331–360 (2014).
Corchón, M. S., Álvarez-Fernández, E. Nuevas evidencias de restos de mamíferos marinos en el Magdaleniense: los datos de La Cueva de Las Caldas (Asturias, España). Munibe 59, 47–66 (2008).
Lefebvre, A. et al. Interconnected Magdalenian societies as revealed by the circulation of whale bone artefacts in the Pyreneo-Cantabrian region. Quat. Sci. Rev. 251, 106692 (2021).
Cortés-Sánchez, M. et al. Earliest known use of marine resources by Neanderthals. PLoS ONE 6, e24026 (2011).
Gutiérrez-Zugasti, I. et al. A chrono-cultural reassessment of the levels VI–XIV from El Cuco rock-shelter: a new sequence for the Late Middle Paleolithic in the Cantabrian region (northern Iberia). Quat. Intel. 474, 44–55 (2018).
Zilhão, J. et al. Last Interglacial Iberian Neandertals as fisher-hunter-gatherers. Science 367, eaaz7943 (2020).
Álvarez-Fernández, E. Marine resource exploitation during the Middle and Early Upper Paleolithic in. Europe: Overv. available Evid. P@lethnology 7, 188–205 (2015).
Fano, M. A., Gutiérrez-Zugasti, I., Álvarez-Fernández, E., Fernández-García, R. Late Glacial and Postglacial use of marine resources in the Bay of Biscay, North Spain. In: Shell energy: Mollusc shells as coastal resources, Bailey, G. N., Hardy, K., Camara, A. eds., Oxbow Books, Oxford, 155–166 (2013).
Aura, J. E. et al. Palaeolithic – Epipalaeolithic Seapeople of the Southern Iberian coast (Spain): an overview. In: Dupont, C., Marchand, G. (eds.), Archéologie des chasseurs-cueilleurs maritimes. De la fonction des habitats à l’organisation de l’espace littoral, Paris, Société préhistorique française, 69–92 (2016).
Bicho, N., Haws, J. A., Davis, L. G. (eds). Trekking the Shore. Changing Coastlines and the Antiquity of Coastal Settlement, New York, Springer, (2011).
Stewart, K., Cunnane, S., Tattersall, I. eds. 2014. The role of freshwater and marine resources in the evolution of the human diet, brain and behavior. Journal of Human Evolution 77, 1–216.
Dupont, C., Marchand, G. (eds). Archéologie des chasseurs-cueilleurs maritimes. De la fonction des habitats à l’organisation de l’espace littoral, Paris, Société préhistorique française, 425 (2016).
Pétillon, J.-M. Circulation of whale-bone artifacts in the northern Pyrenees during the Late Upper Paleolithic. J. Hum. Evolution 65, 525–543 (2013).
Rodrigues, A. S. L., Horwitz, L. K., Monsarrat, S. & Charpentier, A. Ancient whale exploitation in the Mediterranean: species matters. Antiquity 90, 928–938 (2016).
Buckley, M., Collins, M., Thomas-Oates, J. & Wilson, J. C. Species identification by analysis of bone collagen using matrix-assisted laser desorption/ionisation time-of-flight mass spectrometry. Rapid Commun. Mass Spectrom. 23, 3843–3854 (2009).
Collins, M. et al. ZooMS: the collagen barcode and fingerprints. Spectrosc. Eur. 22, 6–10 (2010).
Pétillon, J.-M. et al. Échos de l’océan: phoques et baleines en Europe au Paléolithique récent. In: Cattelain, P., Gillard, M., Smolderen A. (eds.), Disparus? Les mammifères au temps de Cro-Magnon en Europe, Treignes, Cedarc, 335–354 (2018).
Christensen, M. et al. L’industrie osseuse des chasseurs-cueilleurs: le cas des nomades marins de Patagonie et Terre de Feu, Punta Arenas, Universidad de Magallanes. (2016).
Gray, N. M., Kainec, K., Madar, S., Tomko, L. & Wolfe, S. Sink or swim? bone density as a mechanism for buoyancy control in early Cetaceans. Anat. Rec. 290, 638–653 (2007).
Lyman, R. L. et al. Bone density and bone attrition. In: Pokines, J. T., L’Abbe, E. N., Symes, S. A. (eds.), Manual of forensic taphonomy, Boca Raton, CRC Press (2021).
Müller, K. & Reiche, I. Differentiation of archaeological ivory and bone materials by micro- PIXE/PIGE with emphasis on two Upper Palaeolithic key sites: Abri Pataud and Isturitz, France. J. Archaeological Sci. 38, 3234–3243 (2011).
Reiche, I., Müller, K., Staude, A., Goebbels, J. & Riesenmeier, H. Synchrotron radiation and laboratory micro X-ray computed tomography—useful tools for the material identification of prehistoric objects made of ivory, bone or antler. J. Anal. At. Spectrom. 26, 1802–1812 (2011).
Waelbroeck, C. et al. Sea-level and deepwater temperature changes derived from benthic foraminifera isotopic records. Quat. Sci. Rev. 21, 295–305 (2002).
Pétillon, J.-M., Chauvière, F.-X. Les mammifères marins. In: Cretin, C., Madelaine, S. (eds.), Animaux rares, gibiers inattendus: reflets de la biodiversité, Les Eyzies-de-Tayac – Sireuil, Musée national de préhistoire, 92–96 (2019).
Pétillon, J.-M. et al. A gray whale in Magdalenian perigord. Species identification of a bone projectile point from La Madeleine (Dordogne, France) using collagen fingerprinting. Paleo 30, 230–242 (2019).
Lucas, C. et al. The Magdalenian osseous industry from Courbet cave (Penne, Tarn, France) in the British Museum collections. Bull. de. la Soci.été pr.éhistorique française 120, 135–160 (2023).
Langlais, M. et al. Des segments chronoculturels au modèle archéo-stratigraphique du Magdalénien dans le Sud-Ouest français (21 000-16 000 cal. BP). In: Straus, L. G., Langlais, M. eds. Magdalenian chrono-stratigraphic correlations and cultural connections between Cantabrian Spain and Southwest France and beyond, Paris, Société préhistorique française, 109–135 (2020).
Pétillon, J.-M. Technological evolution of hunting implements among Pleistocene hunter-gatherers: osseous projectile points in the Middle and Upper Magdalenian (19–14 ky cal BP). Quat. Int. 414, 108–134 (2016).
Langley, M. C. & Street, M. Long range inland-coastal networks during the Late Magdalenian: evidence for individual acquisition of marine resources at Andernach-Martinsberg. Ger. Cent. Rhineland. J. Hum. Evolution 64, 457–465 (2013).
Laran, S. et al. Seasonal distribution and abundance of cetaceans within French waters – Part II: The Bay of Biscay and the English Channel. Deep Sea Res. Part II: Topical Stud. Oceanogr. 141, 31–40 (2017).
Westley, K. & Dix, J. The solutrean atlantic hypothesis: a view from the ocean. J. North Atl. 1, 85–98 (2008).
Roman, J. et al. Whales as marine ecosystem engineers. Front. Ecol. Environ. 12, 377–385 (2014).
Doughty, C. E. et al. Global nutrient transport in a world of giants. Proc. Natl Acad. Sci. 113, 868–873 (2016).
Gilbert, L., Jeanniard-du-Dot, T., Authier, M., Chouvelon, T. & Spitz, J. Composition of cetacean communities worldwide shapes their contribution to ocean nutrient cycling. Nat. Commun. 14, 5823 (2023).
Wilson, D. E., Mittermeier, R. A. Handbook of the Mammals of the World: Sea Mammals, Barcelona, Lynx Edicions, 4, 614 (2014).
Reeves, R. R., Smith, T. M., Josephson, E. A. Near-annihilation of a species: right whaling in the North Atlantic. In: Kraus, S. D., Rolland, R. M. (eds.), The Urban Whale: North Atlantic Right Whales at the Crossroads, Harvard, Harvard University Press, 39–74 (2007).
Allen, R. C. & Keay, I. Bowhead Whales in the Eastern Arctic, 1611–1911: population reconstruction with historical whaling records. Environ. Hist. 12, 89–113 (2006).
Rodrigues, A. S. L. et al. Forgotten Mediterranean calving grounds of grey and North Atlantic right whales: evidence from Roman archaeological records. Proc. R. Soc. B 285, 20180961 (2018).
Alter, S. E. et al. Climate impacts on transocean dispersal and habitat in gray whales from the Pleistocene to 2100. Mol. Ecol. 24, 1510–1522 (2015).
Hufthammer, A. K., Arntsen, L., Kitchener, A. C. & Buckley, M. Grey whale (Eschrichtius robustus) in Norwegian waters 2000 years ago. Palaeogeogr., Palaeoclimatol., Palaeoecol. 495, 42–47 (2018).
Garrison, E. G., Morgan, G. S., Mcgrath, K., Speller, C. & Cherkinsky, A. Recent dating of extinct Atlantic gray whale fossils (Eschrichtius robustus), Georgia Bight and Florida, western Atlantic Ocean. PeerJ 7, e6381 (2019).
Buss, D. L. et al. Archaeological evidence of resource utilisation of the great whales over the past two millennia: a systematic review protocol. PLoS ONE 18, e0295604 (2023).
García-Vernet, R., Borrell, A., Víkingsson, G., Halldórsson, S. D. & Aguilar, A. Ecological niche partitioning between baleen whales inhabiting Icelandic waters. Prog. Oceanogr. 199, 102690 (2021).
Borrell, A., Abad-Oliva, N., Gómez-Campos, E., Giménez, J. & Aguilar, A. Discrimination of stable isotopes in fin whale tissues and application to diet assessment in cetaceans. Rapid Commun. Mass Spectrom. 26, 1596–1602 (2012).
Blevins, C. et al. Sex- and age-specific migratory strategies of blue whales in the northeast Pacific. Ocean. Front. Mar. Sci. 9, 944918 (2022).
Caraveo-Patiño, J., Hobson, K. A. & Soto, L. A. Feeding ecology of gray whales inferred from stable-carbon and nitrogen isotopic analysis of baleen plates. Hydrobiologia 586, 17–25 (2007).
Ruiz-Cooley, R. I., Gendron, D., Aguíñiga, S., Mesnick, S. & Carriquiry, J. D. Trophic relationships between sperm whales and jumbo squid using stable isotopes of C and N. Mar. Ecol. Prog. Ser. 277, 275–283 (2004).
Reade, H. et al. Nitrogen palaeo-isoscapes: changing spatial gradients of faunal δ 15 N in late Pleistocene and early Holocene Europe. PLoS ONE 18, e0268607 (2023).
Jory, C. et al. Individual and population dietary specialization decline in fin whales during a period of ecosystem shift. Sci. Rep. 11, 17181 (2021).
Seersholm, F. V. et al. DNA evidence of bowhead whale exploitation by Greenlandic Paleo-Inuit 4000 years ago. Nat. Commun. 7, 13389 (2016).
Philippe, M. Un état des connaissances sur la navigation préhistorique en Europe atlantique. Bull. de. la Soci.été pr.éhistorique française 115, 567–597 (2018).
MacLeod, K. et al. Distribution and abundance of fin whales and other baleen whales in the European Atlantic. https://doi.org/10.13140/RG.2.2.19767.52649 (2009).
Speller, C. et al. Barcoding the largest animals on Earth: Ongoing challenges and molecular solutions in the taxonomic identification of ancient Cetaceans. Philos. Trans. R. Soc. Lond., Ser. B, Biol. Sci. 371, 1702 (2016).
Berganza E., Arribas J. L. El entorno físico de las ocupaciones de Santa Catalina. In: Berganza E., Arribas J.L. (eds.), La Cueva de Santa Catalina (Lekeitio): La intervención arqueológica. Restos vegetales, animales y humanos, Bilbao, Kobie serie BAI 4, 367–377 (2014).
Costamagno, S., Théry-Parisot, I., Castel, J.-C., Brugal, J.-P. Combustible ou non? analyse factorielle et modèles explicatifs sur des ossements brûlés paléolithiques. In: Théry-Parisot, I., Costamagno, S., Henry, A. (eds.), Gestion des combustibles au Paléolithique et au Mésolithique: nouveaux outils, nouvelles interprétations, Oxford, Archaeopress, 69–84 (2009).
Berganza E., Arribas J. L. Estructuras de combustión. In: Berganza E., Arribas J.L. (eds.), La Cueva de Santa Catalina (Lekeitio): La intervención arqueológica. Restos vegetales, animales y humanos, Bilbao, Kobie serie BAI 4, 33–48 (2014).
Costamagno, S., Rigaud, J.-P. L’exploitation de la graisse au Paléolithique. In: Costamagno, S. (ed.), Histoire de l’alimentation humaine: entre choix et contraintes, Paris, CTHS, 134–152 (2014).
Berganza, E., Arribas, J. L. Síntesis interpretativa. In: Berganza E., Arribas J.L. (eds.), La cueva de Santa Catalina (Lekeitio, Bizkaia). Industrias líticas y óseas, colorantes y arte mobiliar, Bilbao, Kobie serie BAI 10, 267–285.(2022).
Hussain, S. T. et al. Was the Late Glacial human occupation of northernmost Europe facilitated by whales? New data and perspectives on lithic technology and the paleoecology of the Vendsyssel area, Northern Jutland, Denmark. J. Isl. Coast. Archaeol. 1–29 https://doi.org/10.1080/15564894.2023.2277727 (2024).
Poplin, F. Le bison en os de baleine d’Isturitz (Pyrénées-Atlantiques). Arch.éologie des. Pyrénées Occidentales et. des. Landes 32, 27–28 (2020).
Fiddyment, S. et al. Animal origin of 13th-century uterine vellum revealed using noninvasive peptide fingerprinting. Proc. Natl Acad. Sci. 112, 15066–15071 (2015).
McGrath, K. et al. Identifying archaeological bone via non-destructive ZooMS and the materiality of symbolic expression: Examples from Iroquoian bone points. Sci. Rep. 9, 11027 (2019).
Strohalm, M., Hassman, M., Košata, B. & Kodíček, M. mMass data miner: an open source alternative for mass spectrometric data analysis. Rapid Commun. Mass Spectrom. 22, 905–908 (2008).
Buckley, M. & Collins, M. Collagen survival and its use for species identification in Holocene-lower Pleistocene bone fragments from British archaeological and paleontological sites. Antiqua 1, e1–e7 (2011).
Kirby, D. P., Buckley, M., Promise, E., Trauger, S. A. & Holdcraft, T. R. Identification of collagen-based materials in cultural heritage. Analyst 138, 4849–4858 (2013).
Buckley, M. et al. Species identification of archaeological marine mammals using collagen fingerprinting. J. Archaeological Sci. 41, 631–641 (2014).
Welker, F. et al. Ancient proteins resolve the evolutionary history of Darwin’s South American ungulates. Nature 522, 81–84 (2015).
Coutu, A. N. et al. Palaeoproteomics confirm earliest domesticated sheep in southern Africa ca. 2000 BP. Sci. Rep. 11, 6631 (2021).
Evans, Z., Paskulin, L., Rahemtulla, F. & Speller, C. F. A comparison of minimally-invasive sampling techniques for ZooMS analysis of bone artifacts. J. Archaeological Sci.: Rep. 47, 103738 (2023).
van der Sluis, L. G. et al. Identification and tentative removal of collagen glue in Palaeolithic worked bone objects: implications for ZooMS and radiocarbon dating. Sci. Rep. 13, 22119 (2023).
Longin, R. New method of collagen extraction for radiocarbon dating. Nature 231, 241–242 (1971).
Stafford, T. W. Jr, Duhamel, R. C., Haynes, C. V. Jr & Brendel, K. Isolation of proline and hydroxyproIine from fossil bone. Life Sci. 31, 931–938 (1982).
Stafford, T. W. Jr, Jull, A. J. T., Brendel, K., Duhamel, R. C. & Donahue, D. Study of bone radiocarbon dating accuracy at the University of Arizona NSF accelerator facility for radioisotope analysis. Radiocarbon 29, 24–44 (1987).
Stafford, T. W. Jr., Brendel, K. & Duhamel, R. C. Radiocarbon, 13C and 15N analysis of fossil bone: removal of humates with XAD-2 resin. Geochimica et. Cosmochima Acta 52, 2257–2267 (1988).
van der Sluis, L. G., Zazzo, A., Tombret, O., Thil, F. & Pétillon, J.-M. Testing the use of XAD resin to remove synthetic contamination from archaeological bone prior to radiocarbon dating. Radiocarbon 65, 1160–1175 (2023).
Wacker, L., Christl, M. & Synal, H.-A. Bats: A new tool for AMS data reduction. Nucl. Instrum. Methods Phys. Res. Sect. B Beam Interact. Mater. 268, 976–979 (2010).
Reimer, P. J. et al. INTCAL04 Terrestrial. Radiocarb. Age Calibration, 0-26 Cal. KYR BP, Radiocarb. 46, 1029–1058 (2004).
Brock et al. Current pretreatment methods for AMS radiocarbon dating at the Oxford radiocarbon accelerator unit (ORAU). Radiocarbon 52, 103–112 (2010).
Steier, P., Liebl, J., Kutschera, W., Wild, E. & Golser, R. Preparation methods of μg carbon samples for 14C Measurements. Radiocarbon 59, 803–814 (2017).
Golser, R. & Kutschera, W. Twenty years of VERA: toward a universal facility for accelerator mass spectrometry. Nucl. Phys. N. 27, 29–34 (2017).
Santos, G. M. et al. AMS 14 C sample preparation at the KCCAMS/UCI facility: status report and performance of small samples. Radiocarbon 49, 255–269 (2007).
Beverly, R. K. et al. The keck carbon cycle AMS laboratory, University of California, Irvine: status report. Radiocarbon 52, 301–309 (2010).
DeNiro, M. J. Postmortem preservation and alteration of in vivo bone collagen isotope ratios in relation to palaeodietary reconstruction. Nature 317, 806–809 (1985).
van Klinken, G. J. Bone collagen quality indicators for palaeodietary and radiocarbon measurements. J. Archaeological Sci. 26, 687–695 (1999).
Guiry, E. J. & Szpak, P. Quality control for modern bone collagen stable carbon and nitrogen isotope measurements. Methods Ecol. Evolution 11, 1049–1060 (2020).
Heaton, T. J. et al. Marine20 – The marine radiocarbon age calibration curve (0-55,000 cal BP). Radiocarbon 62, 779–820 (2020).
Dury, J. P. et al. Species-specific reservoir effect estimates: a case study of archaeological marine samples from the Bering Strait. Holocene 32, 1209–1221 (2022).
Bronk Ramsey, C. Bayesian analysis of radiocarbon dates. Radiocarbon 51, 337–360 (2009).
Heaton, T. J. et al. Marine radiocarbon calibration in polar regions: a simple approximate approach using Marine20. Radiocarbon 65, 848–875 (2023).
Beck, L. et al. Marine reservoir effect of spermaceti, a wax obtained from the head of the sperm whale: a first estimation from museum specimens. Radiocarbon 64, 1607–1616 (2022).
Pieńkowski, A., Coulthard, R. D. & Furze, M. F. A. Revised marine reservoir offset (Δ R) values for molluscs and marine mammals from Arctic North America. Boreas 52, 145–167 (2022).
Birkenmajer, K. & Olsson, U. Radiocarbon dating of whale bones from the 17th century whaling sites at Gashamna, Hornsund, South Spitsbergen. Bulletin of the Polish Academy of Sciences. Earth Sci. 46, 109–132 (1998).
England, J., Dyke, A. S., Coulthard, R. D., McNeely, R. & Aitken, A. The exaggerated radiocarbon age of deposit-feeding molluscs in calcareous environments. Boreas 42, 362–373 (2013).
Foote, A. D. et al. Ancient DNA reveals that bowhead whale lineages survived Late Pleistocene climate change and habitat shifts. Nat. Commun. 4, 1677 (2013).
Monge Soares, A. M. et al. Marine radiocarbon reservoir effect in Late Pleistocene and early Holocene coastal waters off Northern Iberia. Radiocarbon 58, 869–883 (2016).
Mangerud, J., Bondevik, S., Gulliksen, S., Hufthammer, A. K. & Høisæter, T. Marine 14C reservoir ages for 19th century whales and molluscs from the North Atlantic. Quat. Sci. Rev. 25, 3228–3245 (2006).
Reimer, R. & Reimer, P. J. An online application for ΔR calculation. Radiocarbon 59, 1623–1627 (2017).
