Thomas, C. D. et al. Extinction risk from climate change. Nature 427, 145–148 (2004).
IPCC. Climate Change 2021: The Physical Science Basis. Contribution of Working Group I to the Sixth Assessment Report of the Intergovernmental Panel on Climate Change. vol. In Press (2021).
Sala, O. E. et al. Global biodiversity scenarios for the year 2100. Science 287, 1770–1774 (2000).
Kazmi, F. A. et al. Ecological impacts of climate change on the snow Leopard (Panthera unica) in South Asia. Braz J. Biol. 82, e240219 (2022).
Aryal, A. et al. Predicting the distributions of predator (snow leopard) and prey (blue sheep) under climate change in the himalaya. Ecol. Evol. 6, 4065–4075 (2016).
Lovari, S. et al. Common and snow leopards share prey, but not habitats: competition avoidance by large predators? J. Zool. 291, 127–135 (2013).
Bocci, A., Lovari, S., Khan, M. Z. & Mori, E. Sympatric snow leopards and Tibetan wolves: coexistence of large carnivores with human-driven potential competition. Eur. J. Wildl. Res. 63, 92 (2017).
Lovari, S., Ventimiglia, M. & Minder, I. Food habits of two Leopard species, competition, climate change and upper treeline: a way to the decrease of an endangered species? Ethol. Ecol. Evol. 25, 305–318 (2013).
Fox, J. L. & Chundawat, R. S. What is a snow Leopard? Behavior and ecology. in Snow Leopards 13–21. https://doi.org/10.1016/B978-0-12-802213-9.00002-X (Elsevier, 2016).
Hacker, C. et al. Regional comparison of snow Leopard (Panthera uncia) diet using DNA metabarcoding. Biodivers. Conserv. 30, 797–817 (2021).
Forrest, J. L. et al. Conservation and climate change: assessing the vulnerability of snow Leopard habitat to treeline shift in the himalaya. Biol. Conserv. 150, 129–135 (2012).
Aryal, A., Brunton, D. & Raubenheimer, D. Impact of climate change on human-wildlife-ecosystem interactions in the Trans-Himalaya region of Nepal. Theor. Appl. Climatol. 115, 517–529 (2014).
Xu, G., Buzzard, P., Blank, D. & Ma, M. Nature reserve in Xinjiang: a snow leopard paradise or refuge for how long? Selevinia 144–148 (2014).
Schaller, G. B., Hong, L., Talipu, Junrang, R. & Mingjiang, Q. The snow Leopard in Xinjiang, China. Oryx 22, 197–204 (1988).
Watts, S. M., McCarthy, T. M. & Namgail, T. Modelling potential habitat for snow leopards (Panthera uncia) in Ladakh, India. PLoS ONE. 14, e0211509 (2019).
Atzeni, L. et al. Meta-replication, sampling bias, and multi‐scale model selection: A case study on snow Leopard (Panthera uncia) in Western China. Ecol. Evol. 10, 7686–7712 (2020).
Din, J. U. et al. On the snow Leopard trails: occupancy pattern and implications for management in the Pamir. Saudi J. Biol. Sci. 29, 197–203 (2022).
Buzzard, P. J., MaMing, R., Turghan, M., Xiong, J. & Zhang, T. Presence of the snow Leopard Panthera uncia confirmed at four sites in the Chinese Tianshan mountains. Oryx 51, 594–596 (2017).
Zhuo, Y. et al. Border fences reduce potential for transboundary migration of Marco Polo sheep (Ovis Ammon polii) in the Pamir plateau. Sci. Total Environ. 912, 169298 (2024).
Huang, D. et al. Biomod2 modeling for predicting the potential ecological distribution of three Fritillaria species under climate change. Sci. Rep. 13, 18801 (2023).
Li, J. et al. Climate refugia of snow leopards in high Asia. Biol. Conserv. 203, 188–196 (2016).
Abedin, I., Singha, H., Kang, H. E., Kim, H. W. & Kundu, S. Forecasting suitable habitats of the clouded Leopard (neofelis nebulosa) in Asia: insights into the present and future climate projections within and beyond extant boundaries. Biology 13, 902 (2024).
Ali, H. et al. Expanding or shrinking? Range shifts in wild ungulates under climate change in Pamir-Karakoram mountains, Pakistan. PLoS ONE. 16, e0260031 (2021).
Mainali, K. P. et al. Projecting future expansion of invasive species: comparing and improving methodologies for species distribution modeling. Global Change Biol. 21, 4464–4480 (2015).
Larson, E. R. & Olden, J. D. Using avatar species to model the potential distribution of emerging invaders. Global Ecol. Biogeogr. 21, 1114–1125 (2012).
Qin, A. et al. Predicting the current and future suitable habitats of the main dietary plants of the gobi bear using maxent modeling. Global Ecol. Conserv. 22, e01032 (2020).
Dyderski, M. K., Paź, S., Frelich, L. E. & Jagodziński, A. M. How much does climate change threaten European forest tree species distributions? Global Change Biol. 24, 1150–1163 (2018).
Takemoto, K. & Kajihara, K. Human impacts and climate change influence nestedness and modularity in Food-Web and mutualistic networks. PLOS ONE. 11, e0157929 (2016).
Dong, H. et al. Climate dry-wet conditions, changes, and their driving factors in Xinjiang. Arid Zone Res. 40, 1875–1884 (2023).
Kramer-Schadt, S. et al. The importance of correcting for sampling bias in maxent species distribution models. Divers. Distrib. 19, 1366–1379 (2013).
Radosavljevic, A. & Anderson, R. P. Making better M axent models of species distributions: complexity, overfitting and evaluation. J. Biogeogr. 41, 629–643 (2014).
Flato, G. et al. Evaluation of climate models. in Climate Change 2013: the Physical Science Basis. Contribution of Working Group I To the Fifth Assessment Report of the Intergovernmental Panel on Climate Change (ed Stocker, T.) F 741–882 (Cambridge University Press, Cambridge, UK, doi:https://doi.org/10.1017/CBO9781107415324.020. (2013).
Running, S. & Zhao, M. MOD17A3HGF MODIS/Terra net primary production Gap-Filled yearly L4 global 500 m SIN grid V006. NASA EOSDIS Land. Processes Distrib. Act. Archive Cent. https://doi.org/10.5067/MODIS/MOD17A3HGF.006 (2019).
Didan, K. MODIS/Terra vegetation indices 16-Day L3 global 1km SIN grid V061. NASA EOSDIS Land. Processes Distrib. Act. Archive Cent. https://doi.org/10.5067/MODIS/MOD13A2.061 (2021).
Wildlife Conservation Society-WCS & Center For International Earth Science Information Network-CIESIN-Columbia University. Last of the Wild Project, Version 2, 2005 (LWP-2): Global Human Influence Index (HII) Dataset (Geographic). Palisades, NY: NASA Socioeconomic Data and Applications Center (SEDAC) (2005). https://doi.org/10.7927/H4BP00QC
Tsendbazar, N. E. et al. Copernicus Global Land Service: Land Cover 100m: Version 3 Globe 2015–2019: Validation Report. https://zenodo.org/record/3606370. https://doi.org/10.5281/ZENODO.3606370 (2021).
Warren, D. & Dinnage, R. ENMTools: Analysis of Niche Evolution Using Niche and Distribution Models. (2024).
Li, J. et al. Defining priorities for global snow Leopard conservation landscapes. Biol. Conserv. 241, 108387 (2020).
Thuiller, W., Lafourcade, B., Engler, R. & Araújo, M. B. BIOMOD – a platform for ensemble forecasting of species distributions. Ecography 32, 369–373 (2009).
Ridgeway, G. GBM: Generalized Boosted Regression Models. R package version 1.6–3.1 (2010).
McCarthy, T. M., Fuller, T. K. & Munkhtsog, B. Movements and activities of snow leopards in Southwestern Mongolia. Biol. Conserv. 124, 527–537 (2005).
VanDerWal, J., Shoo, L. P., Graham, C. & Williams, S. E. Selecting pseudo-absence data for presence-only distribution modeling: how Far should you stray from what you know? Ecol. Model. 220, 589–594 (2009).
Allouche, O., Tsoar, A. & Kadmon, R. Assessing the accuracy of species distribution models: prevalence, kappa and the true skill statistic (TSS). J. Appl. Ecol. 43, 1223–1232 (2006).
McPHERSON, J. M., Jetz, W. & Rogers, D. J. The effects of species’ range sizes on the accuracy of distribution models: ecological phenomenon or statistical artefact? J. Appl. Ecol. 41, 811–823 (2004).
Fielding, A. H. & Bell, J. F. A review of methods for the assessment of prediction errors in conservation presence/absence models. Envir Conserv. 24, 38–49 (1997).
Li, J. et al. Projected impacts of climate change on snow Leopard habitat in Qinghai Province, China. Ecol. Evol. 11, 17202–17218 (2021).
Xue, Y. et al. Assessing the vulnerability and adaptation strategies of wild camel to climate change in the Kumtag desert of China. Global Ecol. Conserv. 29, e01725 (2021).
Li, J. et al. Identifying potential refugia and corridors under climate change: A case study of endangered Sichuan golden monkey (Rhinopithecus roxellana) in Qinling mountains, China. Am. J. Primatol. 80, e22929 (2018).
Thuiller, W., Lavorel, S. & Araújo, M. B. Niche properties and geographical extent as predictors of species sensitivity to climate change. Glob. Ecol. Biogeogr. 14, 347–357 (2005).
McGarigal, K., Cushman, S. A., Neel, M. C. & Ene, E. & others. FRAGSTATS: spatial pattern analysis program for categorical maps. Computer software program produced by the authors at the University of Massachusetts, Amherst. Available at the following web site. www.umass.edu/landeco/research/fragstats/fragstats.html 6. (2002).
Sun, S., Hong, Y., Guo, J., Zhang, N. & Zhang, M. Landscape Dynamics and Ecological Risk Assessment of Cold Temperate Forest Moose Habitat in the Great Khingan Mountains, China. Biology 12, 1122 (2023).
Acharya, K. P., Paudel, P. K., Jnawali, S. R., Neupane, P. R. & Köhl, M. Can forest fragmentation and configuration work as indicators of human–wildlife conflict? Evidences from human death and injury by wildlife attacks in Nepal. Ecol. Indic. 80, 74–83 (2017).
Zhang, L., Dong, T., Xu, W. & Ouyang, Z. Assessment of habitat fragmentation caused by traffic networks and identifying key affected areas to facilitate rare wildlife conservation in China. Wildl. Res. 42, 266–279 (2015).
Zhang Xiaoyu, W. H. & Xiaogang, N. Impact of human footprint on landscape fragmentation in the Northeastern China tiger and Leopard National park. Acta Ecol. Sin. 42, 4688 (2022).
Tang Mingkun, F. Y., Xu, G. Study on landscape pattern characteristics and conservation strategies of giant panda habitat in the Minshan mountains of Sichuan Province. J. Sichuan Sci. Technol. 42, 5–11 (2021).
Johansson, Ö. et al. Land sharing is essential for snow Leopard conservation. Biol. Conserv. 203, 1–7 (2016).
Li, X. et al. Potential range shift of snow Leopard in future climate change scenarios. Sustainability 14, 1115 (2022).
Xu, F. et al. Winter habitat use of snow leopards in Tomur National nature reserve of Xinjiang, Northwest China: winter habitat use of snow leopards in Tomur National nature reserve of Xinjiang, Northwest China. J. Arid Land. 4, 191–195 (2012).
Bai, D. et al. Assessment of habitat suitability of the snow Leopard (Panthera uncia) in Qomolangma National nature reserve based on maxent modeling. Zool. Res. 39, 373–386 (2018).
Johansson, Ö., Nyam, E., Lkhagvajav, P., Shanti Alexander, J. & Samelius, G. Predation Patterns and Hunting Behavior of Snow Leopards: Insights from an Ibex Hunt. SLR 2, (2023).
Hameed, S. et al. Identifying priority landscapes for conservation of snow leopards in Pakistan. PLoS ONE. 15, e0228832 (2020).
Rashid, W. et al. Modelling potential distribution of snow leopards in Pamir, Northern Pakistan: implications for Human–Snow Leopard conflicts. Sustainability 13, 13229 (2021).
Gong, J. et al. MaxEnt modeling for predicting suitable habitats of snow Leopard (Panthera uncia) in the Mid-Eastern Tianshan mountains. J. Resour. Ecol. 14, 1075–1085 (2023).
Rosenbaum, B. et al. Seasonal space use and habitat selection of GPS collared snow leopards (Panthera uncia) in the Mongolian Altai range. PLoS ONE. 18, e0280011 (2023).
Ismaili, R. R. R. et al. Modeling habitat suitability of snow leopards in Yanchiwan National reserve, China. Animals 14, 1938 (2024).
Aryal, A., Brunton, D., Ji, W. & Raubenheimer, D. Blue sheep in the Annapurna conservation area, Nepal: habitat use, population biomass and their contribution to the carrying capacity of snow leopards. Integr. Zool. 9, 34–45 (2014).
Baral, K. et al. Impact of climate change on distribution of common Leopard (Panthera pardus) and its implication on conservation and conflict in Nepal. Heliyon 9, e12807 (2023).
Reshamwala, H. S. et al. Modeling potential impacts of climate change on the distribution of wooly Wolf (canis lupus chanco). Front. Ecol. Evol. 10, 815621 (2022).
Suel, H., Mert, A. & Yalcinkaya, B. Changing potential distribution of Gray Wolf under climate change in lake district, Turkey. Appl. Ecol. Environ. Res. 16, 7129–7137 (2018).
Dar, S. A. et al. Projected climate change threatens Himalayan brown bear habitat more than human land use. Anim. Conserv. 24, 659–676 (2021).
Mahdavi, T., Shams-Esfandabad, B., Toranjzar, H., Abdi, N. & Ahmadi, A. Potential impact of climate change on the distribution of the Eurasian lynx (lynx lynx) in Iran (mammalia: felidae). Zool. Middle East. 66, 107–117 (2020).
Snow Leopard Network. (2014). Snow Leopard Survival Strategy. Revised 2014 Version Snow Leopard Network, Seattle, Washington, USA. (2014).
Wang, G., Li, Y., Wu, Q. & Wang, Y. Impacts of permafrost changes on alpine ecosystem in Qinghai-Tibet plateau. SCI. CHINA SER. D. 49, 1156–1169 (2006).
Alexander, J. S., Zhang, C., Shi, K. & Riordan, P. A spotlight on snow Leopard conservation in China. Integr. Zool. 11, 308–321 (2016).
Li, X. et al. Ongoing threats and the current status of snow Leopard conservation in China. Biodivers. Sci. 27, 932–942 (2019).
Farrington, J. D. & Li, J. Climate change impacts on snow Leopard range. in Snow Leopards 81–93 (Elsevier, 2024).
Flather, C. H., Hayward, G. D., Beissinger, S. R. & Stephens, P. A. Minimum viable populations: is there a ‘magic number’ for conservation practitioners? Trends Ecol. Evol. 26, 307–316 (2011).
Riordan, P., Cushman, S. A., Mallon, D., Shi, K. & Hughes, J. Predicting global population connectivity and targeting conservation action for snow Leopard across its range. Ecography 39, 419–426 (2016).
Johansson, Ö., Alexander, J. S., Lkhagvajav, P., Mishra, C. & Samelius, G. Natal dispersal and exploratory forays through atypical habitat in the mountain-bound snow Leopard. Ecology 105, e4264 (2024).
Liang, G., Li, Y. & Zhou, L. Promoting the conservation of the Leopard in North China based on habitat suitability and connectivity. Global Ecol. Conserv. 50, e02817 (2024).
Wang, Y. et al. Big cats persisting in human-dominated landscape: habitat suitability and connectivity of leopards in central North China. Landsc. Ecol. 39, 94 (2024).
Malla, R. et al. Habitat suitability of four threatened Himalayan species: Asiatic black Bear, common Leopard, musk deer, and snow Leopard. PeerJ 11, e16085 (2023).
Grace, J., Berninger, F. & Nagy, L. Impacts of climate change on the tree line. Ann. Bot. 90, 537–544 (2002).
Feng, X., Li, J., Liu, H. & Zhang, J. Impacts of global warming on the snow leopard: in Proceedings of the 11th International Conference on Biomedical Engineering and Bioinformatics 249–253 (SCITEPRESS – Science and Technology Publications, Tianjin, China (2022).